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Monographia Aquifoliacearum - Loizeau Pierre-André & Gabrielle Barriera


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Aquifoliaceae of Neotropics

Loizeau P.-A. & G. Barriera


Ilex danielis Killip & Cuatrecasas

Habit, vegetative morphology.— Tree or shrub (petit), evergreen, 2–4 m tall. 1 cm dbh. Current year's branchlets glabrous or pubescent. Branchlets older than one year smooth or furrowed, greyish or brownish or reddish, glabrous or pubescent (poils minuscules), sparse trichomes or dense short trichomes. Lenticels on current year's branchlets present. Internodes 0.1–1.6 cm. Stipules caducous, deltoid or linear-triangular, 0.2–0.7 mm. Petiole glabrous or pubescent (poils très courts), smooth surface or rough or grooved surface, 3–12 mm. Lamina elliptic or obovate, oblong (1:2), 2–6.5 X, 1.4–3.7 cm, coriaceous (papyracé chez les très jeunes feuilles), glabrous abaxially (base du limbe pubescent chez les très jeunes feuilles). Base cuneate or acute or obtuse (rarement 3). Apex rounded or obtuse or acute, mucronate or not mucronate. Apex retuse or rounded or acuminate. Acumen 0–0.6 cm. Margin revolute (sauf chez les très jeunes feuilles), entire or dentate, teeth inconspicuous. Teeth 1 ca., on the distal half of the margin. Midbrid adaxially impressed, glabrous (parfois poilue chez les très jeunes feuilles). Midbrid abaxially raised, glabrous (pubescente chez les très jeunes feuilles). Secondary veins abaxially evident or obscure, 4–7 pairs. Tertiary veins abaxially obscure. Ponctuation abaxiallly present (mais très peu visibles) or absent.

Inflorescences.— Thyrse, proliferating.

Male flowers.— Thyrses axes solitary and axillary or terminal. Cymes of order 1–2. Peduncles glabrous or pubescent. Peduncles of order 1 1–10 mm. Pherophylls of order 1 at 4–10 tenth from the base (parfois sur calice). Peduncles of order 2 0.5–2 mm. Pherophylls of order 2 at 2–10 tenth from the base (parfois sur le calice). Bracts linear-triangular, glabrous or pubescent, 0.4–1.5 mm, margin entire or margin ciliate or dentate. Pherophylls of order 1 opposite or subopposite (rarement), deltoid or linear-triangular, glabrous or pubescent, 0.2–0.7 mm, margin entire or margin ciliate. Male flowers 4–5 -merous, white. Calyx glabrous, 1.5–2.5 mm in diam., margin entire or margin ciliate or dentate. Corolla 2–2.5 mm, petals connate on, petals 10–25 % of their length. Apex margin ciliate. Stamens at the anthesis equal to corolla or shorter than corolla. Rudimentary ovary subglobulous or flattened, glabrous.

Female flowers.— Thyrses axes solitary and axillary or terminal. Thyrses axes 0–0.7 cm. Cymes of order 1–2. Peduncles pubescent. Peduncles of order 1 3–13 mm. Pherophylls of order 1 at 2–8 tenth of the base. Peduncles of order 2 3–4 mm. Pherophylls of order 2 at 0–1 tenth of the base. Bracts deltoids or linear-triangulate, pubescent, 0.5–3 mm, margin entire or margin ciliate or dentate. Pherophylls of order 1 opposite or subopposite, deltoids or linear-triangulate, pubescent, 0.3–0.5 mm, margin entire or margin ciliate. Female flowers 4–6 -merous, white. Calyx glabrous, margin entire or margin ciliate or dentate, 1.2–2.8 mm in diam. Corolla 2.5–3 mm, petals free. Apex margin entire (possibly beaded). Staminodes sagittate, glabrous. Ovary spherical, glabrous, stigma capitate, stigma glabrous.

Fruits.— Fruit ovoid or spherical or flattened, 3.5–5 mm in diam., 3–4.5 mm.

Distribution, ecology.— South America, Colombia (Antioquia) Venezuela, 2100–2500 m, Forêt de chêne submontagnarde; arbustal de Bonnetia multinervia (Theaceae).

Flowering.— January or February.


Cite this publication as: ‘Loizeau, P.-A. & G. Barriera (2007). Aquifoliaceae néotropicales: Descriptions, Illustrations, Identification, et Recherche d'Information. Version: 1st march 2007. http://www.ville-ge.ch/cjb/.
Loizeau (1994) et Loizeau & Spichiger (1992) should also be cited (see Bibliography).