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Monographia Aquifoliacearum - Loizeau Pierre-André & Gabrielle Barriera


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Aquifoliaceae of Neotropics

Loizeau P.-A. & G. Barriera


Ilex divaricata Mart. ex Reissek

Habit, vegetative morphology.— Tree or shrub, evergreen, 0.5–5 m tall. Current year's branchlets glabrous. Branchlets older than one year smooth or furrowed, greyish or brownish, glabrous. Lenticels on current year's branchlets present. Internodes 0.2–5 cm. Stipules caducous, linear-triangular, 0.2–0.5 mm. Petiole glabrous, rough or grooved surface (lisse chez les très jeunes feuilles), 4–15 mm. Lamina obovate, oblong (1:2) or lanceolate (1:3), 2.5–11 X, 0.9–5.2 cm, coriaceous, glabrous abaxially. Base cuneate or acute. Apex rounded or obtuse, mucronate or not mucronate. Apex retuse or emarginate or rounded. Margin plane or revolute, entire. Midbrid adaxially flat or raised, glabrous. Midbrid abaxially flat or raised, glabrous. Secondary veins abaxially evident, 5–9 pairs. Tertiary veins abaxially evident or obscure. Ponctuation abaxiallly present.

Inflorescences.— Thyrse, non proliferating or proliferating.

Male flowers.— Thyrses axes solitary and axillary or issued from several axillary buds. Thyrses axes 0.1–0.3 cm. Cymes of order 1–2. Peduncles pubescent. Peduncles of order 1 2–5 mm. Pherophylls of order 1 at 4–9 tenth from the base. Peduncles of order 2 0.1–1.5 mm. Pherophylls of order 2 at 0–1 tenth from the base. Bracts deltoids or linear-triangular, glabrous or pubescent, 0.4–0.8 mm, margin entire or margin ciliate or dentate. Pherophylls of order 1 opposite or subopposite, deltoid, pubescent, 0.2–0.5 mm, margin entire or margin ciliate. Male flowers 4 -merous, white. Calyx pubescent, 1–2.2 mm in diam., margin ciliate or dentate. Corolla 1.5–2.2 mm, petals connate on, petals 5–10 % of their length. Apex margin ciliate. Stamens at the anthesis equal to corolla or shorter than corolla (plus courtes selon LOESENER 1901 et REISSEK 1866). Rudimentary ovary pyramidal, glabrous.

Female flowers.— Thyrses axes solitary and axillary or issued from several axillary buds. Thyrses axes 0.1–0.3 cm. Cymes of order 1. Peduncles glabrous or pubescent. Peduncles of order 1 1–2.5 mm. Pherophylls of order 1 at 6–10 tenth of the base. Bracts deltoids or linear-triangulate, glabrous or pubescent, 0.4–0.8 mm, margin entire or margin ciliate or dentate. Pherophylls of order 1 opposite or subopposite, deltoids or linear-triangulate, glabrous or pubescent, 0.3–0.7 mm, margin entire or margin ciliate. Female flowers 4(–5) -merous, white. Calyx glabrous or pubescent, margin ciliate or dentate, 1.9–3 mm in diam. Corolla 1.5–3.2 mm, petals connate on, petals 5–10 % of their length. Apex margin entire (possibly beaded) or ciliate. Staminodes sagittate, glabrous. Ovary spherical, glabrous, stigma capitate, stigma glabrous.

Fruits.— Fruit green, spherical, 3–5.5 mm in diam., 2–5.5 mm.

Distribution, ecology.— South America, Colombia Venezuela Brazil (Bassin amazonien), 100–750 m, Forêt primaire, bord de savanne.

Illustrations.— • Dessin in Berry & al. (1995). Flora of the Venezuelan Guayana. • Planche Martius s.n. M m. • Rameau Martius s.n. M m. • Pontuations Martius s.n. M m. • Inflorescence Martius s.n. M m. • Inflorescence Martius 3138 M m. • Planche Martius 3138 M m. • Planche Maguire 37646 M f. • Rameau Maguire 37646 M f. • Infrutescence Maguire 37646 M f. • Infrutescence Maguire 37646 M f. • Limbe dessous Maguire 37646 M f. • Planche Maguire 37646 M f. • Rameau Maguire 37646 M f. • Infrutescence, fleur fruit Maguire 37646 M f. • Planche Martius s.n. M m (HT). • Etiquettes Martius s.n. M m (HT). • Rameaux Martius s.n. M m (HT). • Inflorescences Martius s.n. M m (HT).


Cite this publication as: ‘Loizeau, P.-A. & G. Barriera (2007). Aquifoliaceae néotropicales: Descriptions, Illustrations, Identification, et Recherche d'Information. Version: 1st march 2007. http://www.ville-ge.ch/cjb/.
Loizeau (1994) et Loizeau & Spichiger (1992) should also be cited (see Bibliography).