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Monographia Aquifoliacearum - Loizeau Pierre-André & Gabrielle Barriera


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Aquifoliaceae of Neotropics

Loizeau P.-A. & G. Barriera


Ilex nervosa Triana

Habit, vegetative morphology.— Tree or shrub, evergreen, 3–20 m tall. (8–)20–30 cm dbh (pour respectivement des arbres de 12 et 15 m). Current year's branchlets pubescent or hirsute. Branchlets older than one year smooth or furrowed, greyish or brownish or reddish, glabrous. Lenticels on current year's branchlets present or absent or hardly visible. Internodes 0.5–4.8 cm. Stipules persistent, linear-triangular, 1–2 mm. Petiole glabrous or pubescent (par endroit, glabre), smooth surface, 4–18 mm. Lamina ovate to elliptic, orbiculate (1:1) to oblong (1:2), 4–15 X, 2.2–7.5 cm, papyraceous to coriaceous (subcoriace), glabrous abaxially (dessus, quelques poils épars dessous) or pubescent abaxially. Base cuneate to rounded (le limbe se prolongeant parfois le long du pétiole). Apex truncate to acute, not mucronate. Apex retuse or rounded to acuminate. Acumen 0–1.6 cm. Margin revolute (très finement), entire (mais sinueuse) or crenate (finement) or dentate, teeth inconspicuous or conspicuous. Teeth 0–13 ca., over the entire length of the margin or on the distal half of the margin. Midbrid adaxially impressed or flat, glabrous or pubescent on proximal half or pubescent. Midbrid abaxially raised, pubescent on proximal half to pubescent. Secondary veins abaxially evident, 6–13 pairs. Tertiary veins abaxially evident. Ponctuation abaxiallly absent.

Inflorescences.— Thyrse, proliferating.

Male flowers.— Thyrses axes solitary and axillary or issued from several axillary buds (2 méristèmes). Thyrses axes 0.2–0.5 cm. Cymes of order 1–3. Peduncles glabrous or pubescent. Peduncles of order 1 7–14 mm. Pherophylls of order 1 at 8–10 tenth from the base. Peduncles of order 2 0.5–2.5 mm. Pherophylls of order 2 at 2–3 tenth from the base. Peduncles of order 3 0.5–1 mm. Pherophylls of order 3 at 3–5 tenth from the base. Bracts deltoids or linear-triangular, pubescent, 0.5–2 mm, margin entire or margin ciliate or dentate. Pherophylls of order 1 opposite or subopposite, deltoid or linear-triangular, glabrous or pubescent (poils épars), 0.2–0.5 mm, margin entire. Male flowers 4–6 -merous (rarement 4-mère), white. Calyx pubescent, 3–3.5 mm in diam., margin entire or margin ciliate or dentate. Corolla 3–4 mm, petals connate on, petals 10–25 % of their length. Apex margin entire (papilleuse) or ciliate. Stamens at the anthesis equal to corolla or shorter than corolla. Rudimentary ovary pyramidal or subglobulous or flattened, glabrous.

Female flowers.— Thyrses axes solitary and axillary. Thyrses axes 0.5–1 cm. Cymes of order 1–2 (-3 ?). Peduncles glabrous or pubescent. Peduncles of order 1 10–13 mm. Pherophylls of order 1 at 7–8 tenth of the base. Peduncles of order 2 3–4 mm. Pherophylls of order 2 at 0–3 tenth of the base. Bracts deltoids or linear-triangulate, glabrous or pubescent, 0.2–1.2 mm, margin entire or margin ciliate or dentate. Pherophylls of order 1 opposite or subopposite, deltoids or linear-triangulate, glabrous or pubescent, 0.5–1 mm, margin entire or margin ciliate. Female flowers 4–7 -merous, white. Calyx glabrous or pubescent, margin entire (à part quelques poils) or margin ciliate or dentate, 1.5–3 mm in diam. Corolla 2–2.4 mm, petals connate on, petals 20–25 % of their length. Apex margin ciliate. Staminodes sagittate, glabrous. Ovary spherical or flattened, glabrous, stigma mamillate or capitate, stigma glabrous.

Fruits.— Fruit black, spherical, 3–7 mm in diam., 3–7 mm (les fruits semblent contenir 2 ovules par loge).

Distribution, ecology.— South America, Colombia Ecuador Peru, 1800–3000 m, "Bosque" andin de pérennes et décidues: forêt humide de montagne, forêt de brouillard, forêt tempérée humide.

Illustrations.— • Dessin Spruce 5076 G mâle (T du var. aequatoriensis). • Planche Spruce 5076 G m (T du var. aequatoriensis). • Marge limbe Spruce 5076 G m (T du var. aequatoriensis). • Marge limbe Spruce 5076 G m (T du var. aequatoriensis). • Inflorescence Spruce 5076 G m (T du var. aequatoriensis). • Planche Spruce 5076 W m (T du var. aequatoriensis). • Rameau Spruce 5076 W m (T du var. aequatoriensis). • Limbe, face inf. Spruce 5076 W m (T du var. aequatoriensis). • Limbe, face sup. Spruce 5076 W m (T du var. aequatoriensis). • Inflorescence Spruce 5076 W m (T du var. aequatoriensis). • Inflorescence Spruce 5076 W m (T du var. aequatoriensis). • Planche Spruce 5076 NY1. • Planche Spruce 5076 NY2. • Planche Triana 3521(5705.4) BM f (IT). • Limbe, faces supérieure & inférieure Triana 3521(5705.4) BM f (IT). • Infrutescence Triana 3521(5705.4) BM f (IT). • Planche Triana 3521(5705.4) P f (HT). • Limbe face inférieure Triana 3521(5705.4) P f (HT). • Fruits Triana 3521(5705.4) P f (HT). • Infrutescence Triana 3521(5705.4) P f (HT). • Planche Beck 3147 LPB flrs malades. • Etiquette Beck 3147 LPB flrs malades. • Planche Dorr 6710 LPB flrs malades.


Cite this publication as: ‘Loizeau, P.-A. & G. Barriera (2007). Aquifoliaceae néotropicales: Descriptions, Illustrations, Identification, et Recherche d'Information. Version: 1st march 2007. http://www.ville-ge.ch/cjb/.
Loizeau (1994) et Loizeau & Spichiger (1992) should also be cited (see Bibliography).